Media upload via backend doen’t work after moving site

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I have copied my site to a subdirectory for test purposes, everything works fine now. The only problem I have seen so far is the Media upload. When I try to upload a file I get a:

Unable to create directory wp-content/uploads/2016/05. Is its parent directory writable by the server?

The wp-content folder has a 755 permission, the uploads folder also.

I tried to add this to my wp-config

define( 'UPLOADS', 'wp-content/'.'uploads');

in the section before the table prefix is set, because I thought maybe I have to tell WP that the path changed

    define( 'UPLOADS', 'unterverzeichnis/wp-content/'.'uploads');

but I get the same error.

Any Ideas?

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